1 Precipitation in arid regions is temporally variable with much of it arriving in discrete, unpredictable pulses. Climate change models predict an increase in the variation of precipitation, with longer droughts and larger rainfall events, in addition to increased temperatures. 2 A life table response experiment (LTRE) was conducted with the herbaceous arid-land perennial Cryptantha flava (Boraginaceae) from 1997 to 2000, in order to determine how variation in precipitation affects asymptotic population growth (l) and vital rates. Variation in precipitation took two forms, through rainout shelters erected just before and during the 1999 spring growing season, and through naturally occurring variation over the 4 years. 3 An unexpected [...]