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Public Summary: The area burned by wildfires is expected to increase in many watersheds of the world over the next century as a function of climate change. Increased sedimentation due to soil erosion in burned watersheds can negatively impact downstream aquatic ecosystems and the quality and supply of water. At least 65% of the water supply in the western USA originates in watersheds covered by trees, shrubs, and/or grasses that are prone to wildfire16. Understanding how changing fire frequency, extent, and location will affect watersheds, reservoirs, and the ecosystem services they supply to communities is therefore of great societal importance. A primary threat to socio-ecological systems in this region from...